I am a theoretical biologist working on social evolution theory and modelling behavioural evolution more generally.

Research interests
I am interested in how evolution works and shapes complex animal behaviours.

My work has focused on social evolution specifically the evolution of altruistic and spiteful trait in bacteria and multicellular life.

I am also interested in using artificial life and simulations to ask more general questions about co-evolution and cooperation across species.

I have recently also been working on the evolution of intelligence and how we can use major transitions theory to better understand the evolution of animal and non-animal intelligences.