John obtained a BA in Zoology at the University of Cambridge and his PhD from the University of London and the Imperial Cancer Research Fund. After a post-doctoral research period at the MRC National Institute for Medical Research, John was awarded an MRC Career Development Fellowship followed by an MRC Senior Non-Clinical Fellowship at University College London. He joined the Department of Pharmacology, University of Oxford in 2002 and took up a University Lectureship in 2006. John is now an Associate Professor in Molecular and Cellular Pharmacology and a Tutorial Fellow in Medicine at Worcester College, Oxford.