Residual Stand Structure and Topography Predict Initial Survival and Animal Browsing of Redwood and Douglas-Fir Seedlings Planted in Coastal Forests of Northern California

Successful regeneration of commercial species is central to the long-term sustainability of forests managed for wood production. We studied two species of tree seedlings planted after group selection and single-tree selection harvesting in a 20 ha replicated silviculture experiment in stands dominated by coast redwood (Sequoia sempervirens (D.Don) Endl.). Treatments consisted of complete harvest in 1 ha group selection opening (GS), low-density dispersed retention (LD), aggregated high-density retention (HA), and dispersed high-density retention (HD). One year after planting, seedlings planted on a southwest aspect had the lowest survival rate, while northeast aspects had nearly complete survival rates. As expected, redwood had a higher survival rate than coast Douglas-fir (Pseudotsuga menziesii var menziesii (Mirb.) Franco). Survival rates exhibited a rise-peak-fall pattern with stand density, most notably on southwest-facing slopes, ranking LD > HA approximate to HD > GS treatments. Deer browsing of planted seedlings was a pervasive problem where Douglas-fir were preferentially browsed over redwood. In treatments with higher retention densities, browsing was less likely, ranking GS > LD > HA > HD treatments. Further from watercourses at higher elevation, the probability of browsing diminished. Overall, dispersed treatments outperformed aggregated and GS treatments by simultaneously maximizing survival and minimizing browsing of planted seedlings. We did not perform site preparation or herbicide treatment of re-sprouting hardwoods following harvest, and therefore recommend testing the effectiveness of understory vegetation management to enhance seedling survival. Consideration could also be given to planting more seedlings in anticipation of lower survival rates, and/or implementing seedling protection measures when and where heavy browsing is expected.