Distribution of overwintering invertebrates in temperate agroforestry systems: Implications for biodiversity conservation and biological control of crop pests

•Understory vegetation strips are uncropped habitats under the trees in agroforestry.•Understory vegetation strips provide overwintering habitats for many invertebrates.•Crop pests overwinter predominantly in crop alleys.•Granivorous carabids overwinter mostly in understory vegetation strips.•Understory vegetation strips are refugia for non-agrobiont carabids.