In his work on cell metabolism he highlights functional aspects such as control, variability, and optimality. Recent projects concern questions around the economy of the cell: the prediction of protein investments based on computational models of metabolism.

Research interests
Theoretical systems biology
Metabolic models
Resource balance analysis
Economic principles in cell metabolism
Other activities
Member of INRAE's national scientific advisory board
Member of managing board PCI Mathematical and computational biology
Forum Economic Principles in Cell Physiology
Book project Economic Principles in Cell Biology