My research focuses on evolution in structured populations: What are the forces which control the nature and distribution of genotypes in subdivided populations and how does this affect the outcome of other evolutionary processes? These questions and others are addressed in a variety of ways, from theoretical analyses to experimental lab model systems to field research. I have investigated the effects of non-equilibrium population structure in the partitioning of genetic variance among populations. Most models of population structure assume uniform populations at equilibrium, which is most unlike most natural populations. My models have included the effects of extinctions, colonizations, population fission events, unequal population sizes, variable migration rates, and other realistic modifications of the theory.