Honors and Awards
National Science Foundation Graduate Fellow 1970-75
Public Health Service Research Fellow, National Cancer Institute 1975-77
Research Career Development Award, National Cancer Institute 1980-85
Passano Foundation Young Scientist Award 1984
Harrison Howe Award 1984
Guggenheim Fellow 1985-86
Pfizer Award in Enzyme Chemistry 1985
Esquire Magazine Register 1985
Denver Post Westerner of the Year 1986
U. S. Steel Award in Molecular Biology 1987
Member, National Academy of Sciences 1987
Honorary Degree, Doctor of Science, Grinnell College 1987
V. D. Mattia Award 1987
Member, American Academy of Arts and Sciences 1988
Newcombe-Cleveland Award, AAAS (with Arthur J. Zaug) 1988
Heineken Prize, Royal Netherlands Academy of Sciences 1988
Gairdner Foundation International Award 1988
Louisa Gross Horwitz Prize (with Phillip A. Sharp) 1988
Albert Lasker Basic Medical Research Award (with Phillip A. Sharp) 1988
Lewis Rosenstiel Award, Brandeis Univ. (with Sidney Altman) 1989
Warren Triennial Prize, Harvard Medical School (with Joan Steitz) 1989
Nobel Prize in Chemistry 1989
Bonfils-Stanton Award for Science 1990
Honorary Degree, Doctor of Science, Univ. of Chicago 1991
Sir Frederick Gowland Hopkins Medal, British Biochemical Society 1992
Doctor of Science (honorary), Drury College, Springfield, MO, 1994
Ciba Geigy-Drew Award in Biomedical Research 1994
Feodor Lynen Medal 1995
National Medal of Science 1995
Mike Hogg Award, M. D. Anderson Cancer Center, Univ. of Texas 1997
Wright Prize, Harvey Mudd College 1998
Doctor of Science (honorary), Colorado College 1999
Doctor of Science (honorary), Univ. of Maryland Baltimore County 2000
Honorary Degree, Doctor of Science, Williams College 2000
Member, Institute of Medicine 2000
Member, American Philosophical Society 2001
Honorary Degree, Charles University, Prague, Czech Republic 2002
Gregor Mendel Medal, Czech Academy of Sciences 2002
Doctor of Science (honoris causa), Ohio State University 2003
Honorary Member of the NAS, Republic of Korea 2004
Rolf Sammet Prize, Goethe University, Frankfurt 2004
Honorary Doctor of Science, Moscow State University, Russia 2004
Honorary Doctor of Science, University of Vermont 2005
ASBMB Award for Exemplary Contribution to Education 2006
Othmer Gold Medal, Chemical Heritage Foundation 2007
Honorary Doctor of Science, University of Buenos Aires, Argentina 2007
Honorary Doctor of Science, Dartmouth College 2008
Lifetime Achievement Award, RNA Society 2009
Honorary Doctor of Science, Rockefeller University 2009
Lifetime Achievement Award, Colorado Bioscience Association 2009
Honorary Doctor of Science, Watson School of Biological Sciences,
Cold Spring Harbor Laboratories, Cold Spring Harbor, NY 2010
Honorary Doctor of Science, Harvard University, Cambridge, MA 2010
Honorary Membership, Alpha Omega Alpha, Honor Medical Society 2011
Honorary Doctor of Engineering, Colorado School of Mines 2013