Title : Functional consequences of genetic loci associated with intelligence in a 1 meta-analysis of 87 , 740 individuals 2 3

Author Affiliations: 9 1 MRC Social, Genetic and Developmental Psychiatry Centre, Institute of Psychiatry, 10 Psychology & Neuroscience, King’s College London, London SE5 8AF, UK. 11 2 NIHR Biomedical Research Centre for Mental Health, South London and Maudsley 12 NHS Trust, London SE5 8AF, UK. 13 3 Department of Medical Epidemiology and Biostatistics, Karolinska Institutet, SE14 17177 Stockholm, Sweden. 15 4 Department of Complex Trait Genetics, VU University, Center for Neurogenomics 16 and Cognitive Research, Amsterdam, 1081 HV, The Netherlands. 17 5 Department of Child and Adolescent Psychiatry, Erasmus University Medical 18 Center, Rotterdam, the Netherlands 19 6 Laboratory of Molecular Neurobiology, Department of Medical Biochemistry and 20 Biophysics, Karolinska Institutet, SE-17177 Stockholm, Sweden. 21 7 Center for Genomic and Computational Biology, Duke University, Durham, NC 22 27708, USA 23 8 Department of Pediatrics, Medical Genetics Division, Duke University, Durham, NC 24 27708, USA 25 9 Departments of Genetics, University of North Carolina, Chapel Hill, NC, 2759926 7264, USA. 27 10 Department of Clinical Genetics, VU University Medical Center (VUMC), 28 Neuroscience Campus Amsterdam, Amsterdam, 1081 HV, The Netherlands. 29 ¶ These authors contributed equally to the work 30